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Toledo is more recent than App State if that takes the sting away
That sucked, but we should have lost to UNI in 2009. It took 2 blocked FGs in the final 5 seconds or so to stop that upset.2016
Iowa lost to North Dakota State (only time Iowa has lost to FCS/DI-AA. Interesting, that same year, Iowa beat #2 (Game day rankings) Michigan.